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The second 43.7 km (27.2 mi) system passes through the centre of Toronto and ends in Pickering to the east.
Progressing eastward, eight lanes are carried beneath the large spaghetti junction at Highway 427.
To avoid confusion between carriageways, blue signs are used for the collector lanes and green signs for the express lanes.
Unlike the collector lanes, which provide access to every interchange, the express lanes only provide direct access to a select few interchanges.
In 1964, the route became fully navigable from Windsor to the Ontario–Quebec border.This area is not as flat but the highway is generally straight.This part of Highway 401 often experiences heavy snowsqualls in early winter, sometimes extending as far east as Toronto.is a controlled-access 400-series highway in the Canadian province of Ontario.It stretches 828.0 kilometres (514.5 mi) from Windsor in the west to the Ontario–Quebec border in the east.
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The highway curves northeast and follows a power transmission corridor to Highway 409, which merges with the mainline and forms the collector lanes.